故事承接《鬼灭之刃柱训练篇》。
Based on the novel by Karel Capek, the prominent Czech writer of the early 20th century who coined the word robot for his play R.U.R., the story revolves around a discovery of Krakatit - a powerful explosive - by Prokop, an experimental scientist who, following an accidental explosion in his lab, slips in and out of delirium. When he realises that, in a delirious period, he has given the formula for making Krakatit to one of his colleagues, he tries to find the man, who does not realize how dangerous a thing Krakatit is. In the process, he nearly gives his discovery away again to foreign interests, a group of anarchists, and even greater forces of darkness...
It's the land we love and the land we think we know. We see America's breathtaking landscapes and wildlife as timeless, but the truth is very different. Its unique geography drives the forces of nature to extremes, shaping and reshaping the land and throwing down new challenges for life.